Coulaceae Tiegh.

Family Description

Distribution Map

Photographs

Coula

1. Coula edulis
   • Habit of small tree and flowering branches.  Loko, Equateur, Democratic Rebublic of Congo. Photos by Roy Danforth, Sept. 26, 2012.  Link goes to Flickr.
   • Herbarium specimen, and close-up of inflorescence Missouri Botanical Gardens (MO). Photo by D. L. Nickrent.
   • Fruits. From a book on Equatorial Guinea plants: Chris Wilks & Yves Issembe. 2001, Practical Identification Guide - Equatorial Guinea Trees - Continental Region. CUREF Publishers.
   • Nuts, photo from a web site of a global nut trader.
   • Shells of nuts cracked open and eaten by humans.  Rainforest preserve at Lake Tissongo, Cameroun.  From Humans and their Environments (University of Pennsylvania): A day in the life of an African hunter.  Photo by D. H. Janzen.
   • Illustration.  From Wikipedia.
   • Article about the reproductive ecology of Coula edulis by Moupela et al. (2014. Tropical Ecology 55:327-338) HERE.
     

Minquartia

1. Minquartia guianensis
   • Series of photos from Atrium, biodiversity information system.
   • Photos. Pasco, Oxapampa, Peru. Link goes to Flora of the World.
   • Series of photos. Vegetative features.  Link goes to PlantSystematics.org.
     
   • Young plant being cultivated at the R. & C. Wilson Botanical Garden, San Vito, Costa Rica. The species is locally called "manu negro" and has very hard, termite-resistant wood. The wood is therefore often used as support beams for buildings. [DLN 2758]. Photo by D. L. Nickrent.
   • Photo of the trunk of a large tree. Costa Rica.  Photo by D. L. Nickrent.  Link goes to PhytoImages.
   • Trunk of tree. Panama. From the Center for Tropical Forest Science, Panama web atlas, Trees of the Panama Canal.
   • Close-up of leaves and buds. Panama. From the Center for Tropical Forest Science, Panama web atlas, Trees of the Panama Canal.
   • Leaf, leaf, close-up, showing characteristic tertiary venation, petiole, showing basal swelling and channel, and broken petiole showing latex in sap. Costa Rica, Heredia La Selva. Photos 2004 by Lawrence Kelly, 2004.
   • Close-up of flowers. Saul, French Guiana. Photo by Carol Gracie.
   • Herbarium specimen photo 1, photo 2, Missouri Botanical Gardens (MO).
     

Ochanostachys

1. Ochanostachys amentacea
   
   • Tree photographed up into its canopy.  From a blog in Singapore.
   • Tree (called petaling) in vegetative condition. Rimbun Dahan, outside Kuala Lumpur, Malaysia. Photos by Hijjas and Angela Kasturi. 
     
   • Bark of tree, shoot with inflorescences and close-up of flowers.  From website called EARTH by Joseph Lai in Singapore.
     
   • Plant with fruits. Malaysia. Photo by A. Gentry (No. 66868). Link goes to TROPICOS.
   • Photos. Leafy branches, leaves, bark features. Link goes to Flickr.
     
   • Illustration. Flora Malesiana 10 12 1984. Link goes to Borneo Tree Images. James K. Jarvie, Beth Dart, Diane Holland.

Phylogeny

• The first molecular phylogeny of Olacaceae s. lat. was by Malécot and Nickrent (2008).  A pdf file for that paper can be seen by clicking HERE.
• A reclassification of the families of Santalales was by Nickrent et al. (2010). A pdf file for that paper canbe seen by clicking HERE.
• A multigene phylogeny for the order Santalales was by Su et al. (2015). A pdf file for that paper canbe seen by clicking HERE.

References